Ricotia Descriptions

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Ricotia in Antalya : Botanical Descriptions

This is an appendix to the Ricotia ID Page primarily is to give some original descriptions for Ricotia since later descriptions are rather brief, the latin can be put into an appropriate web translation service or AI bot, but a rough English version has been provided "as is" from Google Translate which at some point I might correct. "⊙" signifies annual.

Their sequence is: ANNUALS carnosula, sinuata, tenuifolia, cretica, PERENNIALS, candiriana, davisiana, varians.

Ricotia carnosula

1965 Flora Of Turkey

Differs from R. sinuata in its petals (10-)11-12 mm, and linear-oblong, scarcely tapered, up to 12-seeded fruits. Fl. 3-4. Rocky outcrops, s.l.-700 m.

1854 Boissier Diagnoses plantarum Orientalium novarum - Series 1

GOOGLE TRANSLATE: R. small, glaucous, stems are small from the base, very ramified and hairy, fleshy leaves are primordial, ovate, subsessile, subrectangular, red, pinnate or bipinnately divided, with thick margins, minute lower leaves, oblong, upper leaves, elongate, narrowly linear, all obtuse, flowers are white or pink, limb is obcordate, filaments are larger, anthers are barely sesquilonger, anther loculi are subdivergent at the base, pendulous pods are recurved, they are four times shorter, linear, their width is 8-10, and they are longer than 12-15, with subthickened margins, straight or irregularly strangulated, acute, style very short, apiculate. Habitat among sea rocks to the west of the city of Adalia in Pamphylia (Heldr.) FI. Mart, fruct. April. Plant 3-6 inches tall, the leaf margins smaller than those of other species of the genus, especially the upper ones, very narrow. Pods 15-20 lin. long 2 wide.

ORIG: R. pumila glaucescens, caulibus pumilis a basi divaricatim ramosissimis petiolisque hirtulis, foliis carnosulis primordialibus ovatis subsessilibus subretusis rubellis caeteris pinnatim vel bipinnatisectis laciniis margine crassiusculis minutis foliorum inferiorum oblongis superiorum elongatis anguste linearibus omnibus obtusis, floribus albis vel roseis, limbo obcordato, filamentis majoribus anthera vix sesquilon-gioribus, loculis antherae basi subdivergentibus, siliquis pendulis pedicello recurvo eis quadruplo breviori suffultis linearibus latitudine sua 8-10 pio longioribus 12-15 spermis margine subincrassatis rectis aut irregulariter strangulatis acutis stylo brevissimo apiculatis. Hab. inter saxa maritima ad occidentem urbis Adalia in Pamphylia (Heldr.) Fl. Mart, fruct. Aprili. Planta 3-6 pollicaris, laciniae foliorum eis aliarum specierum generis minores superiores praesertim angustissimae. Siliqua 15-20 lin. longa 2 lata.

1867 Flora Orientalis

GOOGLE TRANSLATE: glaucescens, stems small from the base divaricatim ramosis, fleshy leaves pinnately or bipinnately cut segments densely packed, lower leaves ovately lobed, upper narrowly linear, petals obcordate, filaments longer anthers barely sesquilongioribus, anther loculi divergent at the base, siliquis pedicel fourfold longer linear latit. sua 8-10 times longer 12-15 spermis margin straighter acute, style very short ⊙. Hab. among sea rocks near Adalia Pamphyliae (Pestal! Heldr! Bourg!). Fl. Apr. Mai. Plant 4-8 pollicaris, flowers whitish or pink cis R. Lunaria minores, siliqua 15-20 lines long 2 wide.

ORIG: glaucescens, caulibus pumilis a basi divaricatim ramosis, foliis carnosulis pinnatim vel bipinnatim sectis segmentis confertis foliorum infimorum ovatis lobatis superiorum anguste linearibus, petalis obcordatis, filamentis longioribus anthera vix sesquilongioribus, antherae loculis basi divergentibus, siliquis pedicello suo quadruplo longioribus linearibus latit. sua 8-10 plo longioribus 12-15 spermis margine rectiusculis acutis, stylo brevissimo ⊙. Hab. inter saxa maritima prope Adalia Pamphyliae (Pestal! Heldr! Bourg!). Fl. Apr. Mai. Planta 4-8 pollicaris, flores albidi vel rosei cis R. Lunaria minores, siliqua 15-20 lineas longa 2 lata.

Ricotia sinuata

1965 Flora of Turkey

Glabrous annual herb. Leaves pinnate, the segments trifid or pinnatisect, linear, linear-oblong or ovate. Petals white to pale lilac, 7-9(-10) mm, deeply emarginate. Siliquae linear-lanceolate, tapered to the apex, 7-8-seeded, borne on defiexed pedicels. Fl. 4. Rocky slopes, cliffs, etc., s.l-1000 m.

1854 Boissier Diagnoses plantarum Orientalium novarum - Series 1

GOOGLE TRANSLATE:... R. tender, very fragile, with erect stems and short hairy petioles, branched above, loose racemes, bipinnately divided leaves, deeply and obtusely 2-3 lobed petiolate segments, small white flowers, shortly obcordate limb, linear anther filaments, barely longer, subdivergent locules at base, recurved pedicel, six times shorter, linear in width, 7-9 times longer, 7-8 sperms between seeds, strangulated-sinuated, obtuse, rarely acute, style three times shorter, terminating. Habitat: in the jaws of Mount Climax near Kourmalu in Lycia and on the promontory of Alaya in Pamphylia in rocks among shrubs. (Heldreich.) Flowers of R. lunariae are about half smaller, 1 1/2 2 inch long, 2 1/2 lin. wide. It is distinguished from the preceding species [carnosula] by its more tender leaves, more dissected, rounded lobes, and wider, sinuous pods. The three species described here form a new section in the genus by the presence of septa and linear pods, which are striking and very well consistent with the older species, but in habit and characters. Ricotia thus extended differs from Lunaria, however, in its pods not pedicellated, and from Pteroneuro, in its seeds not having any petals or winged filaments.

ORIG: R. tenera fragillima, caulibus erectis petiolisque breviter hirtellis ramosis superne racemos laxos edentibus, foliis bipinnatisectis partitionibus petiolatis segmentis ovato-cuneatis profunde et obtuse 2-3 lobis, floribus parvis albis, limbo breviter obcordato, filamentis anthera lineari vix longioribus, loculis basi subdivergentibus, siliquis pedicello recurvo eis sexluplo breviori suffultis linearibus latitudine sua 7-9 plo longioribus 7-8 spermis inter semina strangulato-sinuatis obtusiusculis rarius acutis stylo latitudine sua triplo breviori terminatis. Hab. in faucibus montis Climax prope Kourmalu Lyciae et in promontorio Alaya Pamphyliae in saxis inter frutices. (Heldreich.) Flores eis R. lunariae fere dimidio minores, siliqua 1 1/2 2 poll. longa 2 1/2 lin. lata. A specie praecedenti [carnosula] distincta foliis tenerioribus magis dissectis lobis rotundatis, siliquis latioribus margine sinuatis. Species ternae hic descriptae in genere novam sectionem formant septi presentia siliquisque linearibus insignem cum specie antiquiori autem habitu et characteribus optime congruentem. Ricotia sic extensa a Lunaria tamen silique non pedicellata seminibus immarginatis, a Pteroneuro nec placentis nec filamentis alatis optime differt.

1867 Flora Orientalis

GOOGLE TRANSLATE:... sparsely hairy, stems decumbent at base, leaves bipinnately divided, segments petiolulated, ovate-cuneate, deeply lobed, upper leaves linear, pelage retuse, filaments anther linear, scarcely longer than the anther loculi divergent at base, siliquas pedicel six times longer than acute, latit. sua 7-9 times longer than 7-8 spermis inter semina strangulate-sinuated, stylus siliquae latit. subtriple shorter ⊙. Habitat. in saxony at promontory Climax Lyciae et prom. Alaya Pamphyliae (Heldi!). in Cilicia Kurdica ad castellum Schech Meran (Ky exs. Suppl. 18!), circa Beilan Syriae borealis (Haussk!). The flowers are white or pink, generally smaller than those of the preceding species, the pods 1 1/2-2 inches long and 2 1/2 lines wide. It differs from R. carnosula in having more dissected leaves, a wider pod with a longer style, and much less blunt petals.

ORIG: R. sinuata (Boiss. et Heldr. Diagn. Ser. I, VIII, p. 30) parce hirtula, caulibus basi decumbentibus, foliis bipinnatisectis segmentis petiolulatis ovato-cuneatis profunde lobatis, foliorum superiorum linearibus, pelalis retusis, filamentis anthera lineari vix longioribus loculis antherae basi divergentibus, siliquis pedicello sextuplo longioribus acutis latit. sua 7-9 plo longioribus 7-8 spermis inter semina strangulato-sinuatis, stylo siliquae latit. subtriplo breviori ⊙. Hab. in saxosis ad promontorium Climax Lyciae et prom. Alaya Pamphyliae (Heldi!). in Cilicia Kurdica ad castellum Schech Meran (Ky exs. Suppl. 18!), circa Beilan Syriae borealis (Haussk!). Flores albi vel rosei eis praecedentis fere minores, siliqua 1 1/2-2 pollicaris 2 1/2 lineas lata. A R. carnosula differt foliis magis dissectis, siliqua latiori longius stylosa, petalis multo minus retusis.

Ricotia tenuifolia

1806 Flora Graeca

GOOGLE TRANSLATE: Root slender, annual, simple; fibrillose at the base. Stem erect, slender, smooth, glabrous, branched on all sides, open, foliate, multiflorous. Leaves alternate, long-petiolate, pinnatifiform, or bipinnatifiform, with opposite, linear, acute, glabrous, subcarnose lobes. Cotyledons obovate, obtuse, very long-petiolate, glabrous, often persistent. Terminal corymbia, solitary, glabrous. Flowers pale fleshy. Calyx glabrous, barely gibbous at the base. Petals obovate, slightly emarginate, with a purple central line; white, narrow nail, the length of the calyx. Stamens all simple, subulate, curved; four longer ones slightly exceeding the bud. Anthers all oblong, erect. Style very short. Silices racemose, pendulous, elliptical, very compressed, glabrous. Seed solitary, kidney-shaped, compressed, reddish, with a very narrow, pale margin, as can be seen from the icon. A doubtful species of this genus.

ORIG: In Lycia. ⊙. Radix gracilis, annua, simplex; inferne fibrillosa. Caulis erectus, gracilis, teres, glaberrimus, undique ramosus, patens, foliosus, multiflorus. Folia alterna, longius petiolata, pinnatificla, vel bipinnatifida, laciniis oppositis, linearibus, acutis, glabris, subcarnosis. Cotyledones obovatae, obtusae, longissime petiolatae, glabrae, saepe persistentes. Corymbi terminales, solitarii, glabri. Flores pallide incarnati. Calyx glaberrimus, vix basi gibbus. Petala obovata, parum emarginata, linea centrali purpurea; ungue albo, angusto, calycis longitudine. Stamina omnia simplicia, subulata, incurva; quatuor longiora germen paululum superantia. Antherae omnes oblongae, erectae. Stylus brevissimus. Siliculae racemosae, pendulae, ellipticae, valde compressae, glabrae. Semen solitarium, reniforme, compressum, rufescens, margine, ut ex icone videtur, angustissimo, pallido. Hujusce generis dubia species.

1867 Flora Orientalis

GOOGLE TRANSLATE:... glabrous erect, with pinnate and bipinnate leaves, with short linear segments, entire, subintact petals, loose fruit clusters, with equilong pedicel, elliptical, obtuse, monospermous, sessile stigma (.). Peltaria Caramaniensis Sibth. in herb. Bancks. Hab. in Lycia (Sibth.). Flowers pale fleshy, preceding them smaller, with 4-5 long lines of silicules (not seen).

ORIG: R. tenuifolia (Sibth. Fl. Graec. tab. 630) glabra erecta, foliis pinnatim et bipinnatim sectis segmentis brevibus linearibus integris, petalis subintegris, racemis fructiferis laxiusculis, siliculis pedicello aequilongis ellipticis obtusis monospermis stigmate sessili (.). Peltaria Caramaniensis Sibth. in herb. Bancks. Hab. in Lycia (Sibth.). Flores pallide incarnati eis praecedentis minores, siliculae 4-5 lineas Iongae (non vidi).

1965 Flora of Turkey

Glabrous annual herb. Leaves 1-2 x pinnate, the segments short, entire, linear. Petals pale red. Siliculae oval, 1-5-seeded [appears to be an error for 1 seeded].

Type: Lycia, Sibrhorp (holo. OXF; iso. BM!).

C3 Antalya: d. Finike, Finike to Elmali, 600 m, Hub.-Mor. 9886.

Endemic; close to R lunaria (L.) DC., from W. Syria and the Syrian Desert.

Ricotia cretica

2016 Atlas of the Aegean

Glabrous annual. Stems several, slender, ascending, 10-30 cm. Basal leaves bright green, ternate or pinnate; leaflets obovate, entire or obtusely lobed. Cauline leaves small. Racemes lax, usually 3-10-flowered, ± secund in fruit. Petals 10-12 mm; limb obovate, deeply emarginate, lilac to bright rose-pink with yellowish base. Siliculae pendent, strongly compressed, oblong, 25-45 x 7-9 mm, not winged, brownish. Seeds 3-5 in each loculus.

1993 Flora Europaea ed 2 vol 1

Annual 10-25 cm. Leaves 2-pinnatifid or -pinnatisect, the upper with ovate or elliptical segments. Petals 10-12 mm, pink or violet. Siliqua 30-50 x 8-9 mm, 5-8 times as long as wide, not winged. Seeds up to 10. Calcareous screes and stony ground. Kriti. Cr.

1867 Flora Orientalis

GOOGLE TRANSLATE:... glabrous, stems decumbent at the base, leaves bipinnately segmented, petiolate, oblong, lobed, petals deeply obcordate, filaments larger than the anthers linear, scarcely sesquilongioribus, siliquis pedicel twice as long as wide, linear latit. sua 7-8 times longer, spermis margine rectiusculis, stylo siliquae latit. quadruple shorter ⊙.

Hab. in the faucibus Askyphus montium Sphacioticorum Cretae (Heldr!), in saxosis maritimis ad Hagios Paulos (Raulin!).

Facies et flores rosei R. Lunarioe. Siliquae 16-25 lines long 3 latae.

ORIG: R. Cretica (Boiss. et Heldr. Diagn. Ser. I, VIII, p. 29) glabriuscula, caulibus basi decumbentibus, foliis bipinnatisectis segmentis petiolatis oblongis lobatis, petalis profunde obcordatis, filamentis majoribus anthera lineari vix sesquilongioribus, siliquis pedicello duplo longioribus late linearibus latit. sua 7-8 plo longioribus 7-9 spermis margine rectiusculis, stylo siliquae latit. quadruplo breviori ⊙. Hab. in faucibus Askyphous montium Sphacioticorum Cretae (Heldr!), in saxosis maritimis ad Hagios Paulos (Raulin!). Facies et flores rosei R. Lunarioe. Siliquae 16-25 lineas longae 3 latae.

1854 Boissier Diagnoses plantarum Orientalium novarum - Series 1

GOOGLE TRANSLATE:... glabrous, tender stems ascending, elongate, foliose, branching into loose racemes, primordial leaves ovate, undivided, the rest bipinnately divided with oblong, obtuse, entire or lobed lacini, the tops trisected with oblong-linear lacini, large pink flowers, edged, filaments larger, linear anthers scarcely sesquilongioribus, pendulous pods with a pedicel half shorter than them, broadly linear in width 7-8, longer than 7-9, sperms with straight, acute margins, style four times shorter than its width, apiculate.

Habit. among the rocks of the Sphaciotic mountains in the mouth of the Askyphous mountain range, said alt. 3000’. (Heldreich) Fl. Aprili.

Stems 3/4-1 pedal, leaves similar to R. lunarioe, flowers almost larger, siliqua 16-20 lin. long 3 wide.

ORIG: R. glabriuscula tenera caulibus ascendentibus elongatis foliosis ramosiss in racemos laxos abeuntibus, foliis primordialibus ovatis indivisis, caeteris bipinnatisectis laciniis oblongis obtusis integris vel lobatis, summis trisectis laciniis oblongo-linearibus integris, floribus roseis magnis, limbo obordato, filamentis majoribus anthera lineari vix sesquilongioribus, siliquis pendulis pedicello eis dimidio breviori suffultis lato-linearibus latitudine sua 7-8 pio longioribus 7-9 spermis margine rectiusculis acutis stylo latitudine sui quadruplo breviori apiculatis. Hab. inter saxa montium Sphacioticorum in fauce Askyphous dicta alt. 3000’. (Heldreich) Fl. Aprili. Caules 3/4-1 pedales, folia eis R. lunarioe similia, flores eis ejus fere majores, siliqua 16-20 lin. longa 3 lata.

Ricotia candiriana

2019 PDF

Perennial, glaucous herbs up to 6.5–18 cm, pilose, especially towards the base. Stem 6–13.5 cm, simply branched, ascending. Leaves decreasing in size from the base to the upper part of the stem; basal leaves pinnate, 26–60 x 7.0–24 mm, generally with 3–5 obovate-lanceolate to lanceolate segments, glabrescent to sparsely pilose, fleshy, green or reddish tinged; terminal segment 9.0–17.0 x 3.0–4.5 mm, entire or 2–3 lobed; lateral segments 7.0–11.0 x 1.5–3.0 mm, usually entire, sometimes upper lateral segments 2-lobed; petiole 15–35 mm. Stem leaves usually pinnate with 3–5 lanceolate to linear-lanceolate segments, sometimes simple, 6–25 x 2.3–14 mm, pilose, segments entire or 2- lobed; terminal segment 4–10 x 1–2 mm, linear-lanceolate to linear; lateral segments 3–9 x 1–2 mm, linear-lanceolate to linear; petiole 3–13 mm. Inflorescence usually with 5–10 flowers, elongating in fruit, pedunculate (peduncle up to 2.5 cm). Flowers pedicellate; pedicels 2–10 mm long; fruiting pedicels much thicker and recurved. Sepals 4.5–5.7 x 0.7–1.3 mm, green or purple tinged, closed, outer sepals slightly saccate at the base, membranous at margins; teeth triangular, 0.75–0.9 x 0.7–0.8 mm, generally purple tinged at the apex, obtuse. Petal claws 3.8–5.4 x 0.35–0.6 mm; lamina 1.4–3.5 x 1.2–2.5 mm, emarginate, mauve, yellowish at the base. Filaments 1.0–3.0 x 0.15–0.32 mm, widened at the base; anthers 1.1–2.7 x 0.2–0.3 mm. Pistil 2.24–2.69 mm long; ovary densely pubescent; stigma capitate. Siliqua 25–60 x 4.0–9.0 mm, sparsely pilose, with marginal veins; septum membranous. Seeds 9–17, winged, broadly elliptic to orbicular, 4.0–6.0 x 3.0–4.0 mm (including wings), light brown, with reticulate ornamentation; wings membranous, 0.4–1.2 mm wide. Endemic... observed to flower in May and June, with mature fruit in June... Kumluca, upper part of Alakır Valley, close to the Dibek Nature Conservation Area, screes, 1160 m elevation. (PDF with photos).

Ricotia davisiana

1965 Flora of Turkey

R. davisiana B. L. Burtt in Kew Bull. 1951: 130 (1952).

Perennial herb, with a dense indumentum of short, simple hairs. Leaves fleshy, glaucous, trifoliolate, the segments elliptic to orbicular, obtuse. Petals lilac-pink. Siliquae linear-oblong, 6-12-seeded. Fl. 7. Limestone serees, 2000-2300 m.

Type: Turkey [C3] vil. Antalya, Tahtali Dağ above Kemer, mobile N screes, 2000-2300 m, Davis 14123 (holo. K!; iso. E!).

Endemic and known only from Tahtali Da. E. Medit. (mt.) element. Related to the annual R. cretica Boiss, & Heldr., endemic to Crete, and to R. varians.

1951 Genus Ricotia

GOOGLE TRANSLATE:... R. creticae is most similar in fruit and seed form, but differs greatly in perennial habit and clothing and leaves always deeply trilobed. Perennial herb, up to 10 cm. high, with many erect-flexible branched stems, with open roots and (in cultivation) emitting innovations; the whole plant is covered with short simple straight open hairs. Fleshy leaves, all long-petiolate, lamina trilobate at the base; petiole 3-4 cm. long; lobes more or less equal (but lateral ones slightly asymmetrical at the base), obovate, rounded or emarginate, entire or irregularly and obtusely lobulated, 1-1.5 cm. long, 8-12 mm. wide; sometimes (in cultivation) a solitary lobe is also produced from the middle of the petiole. Inflorescence terminal, on a bare peduncle about 6 cm. long; flowers about 6, ebracteate. Sepals 4, contiguous, the outer ones slightly saccate at the base, abruptly narrowed at the apex, obtuse, slightly membranous-marginate, briefly and openly pubescent. Petals 4, claw 6 mm. long and 1 mm. wide; very pale pink-purple lamina straight and clearly obcordate at the base, very slightly auriculate at the apex, 6-5 mm. long and 7 mm. wide. Stamens 6; 4 longer filaments 4 mm. long slightly dilated towards the base; shorter filaments 3 mm. long at the base by 0.5 mm. very externally curved; anthers 2 mm. long. Gynoecium 2.25 mm. long, briefly and openly hairy, crowned with a capitate stigma. Fruit pedicel 8 mm. long, supported, very compressed, more or less linear, about 5 cm. long and 6 mm. lateral, clearly pubescent, endowed with a marginal vein and a membranous septum. Seeds in full fruit about 12, disciform, embryo 3 mm. in diameter, seeds hyaline wing 1.5 mm. wide circumscribed; cotyledons 4 mm. long and 2.5 mm. wide, slightly emarginate.

ORIG: R. davisiana B. L. Burtt, sp. nov. R. creticae Boiss. et Heldr. ad formam fructus et seminum maxime affinis, sed habitu perenni et indumento et foliis semper profunde trilobatis valde discrepat. Herba perennis, 10 cm. usque alta, caulibus pluribus ramosis erecto-flexuosis, radicibus patentibus et (in culta) innovationes emittentibus; tota planta pilis brevibus simplicibus recte patentibus vestitis. Folia carnosula, omnia longe petiolata, lamina ad basin trilobata; petiolus 3-4 cm. longus; lobi plus minusve aequales (sed laterales basi leviter asymmetrici), obovati, rotundati vel emarginati, integri vel irregulariter et obtuse lobulati, 1-1.5 cm. longi, 8-12 mm. lati; interdum (in planta culta) lobus solitarius e medio petiolo etiam productus est. Inflorescentia terminalis, pedunculo nudo circiter 6 cm. longo; flores circiter 6, ebracteati. Sepala 4 inter se cohaerentia, exteriores basi leviter saccata, apice abrupte angustata, obtusa, leviter membranaceo-marginata, breviter et patente pubescens. Petala 4, ungue 6 mm. longo et 1 mm. lato; lamina pallidissime roseo-purpurea recte patente obcordata, basi levissime auriculata, apice ad 0.75 mm. emarginata, 6-5 mm. longa et 7 mm. lata. Stamina 6; 4 longiora filamentis 4 mm. longis basin versus leviter dilatata; breviora filamentis 3 mm. longis basi per 0.5 mm. valde externe curvatis; antherae 2 mm. longae. Gynoecium 2.25 mm. longum, breviter et patenter pilosum, stigmate capitato coronatum. Fructus pedicello 8 mm. longo suffultus, valde compressus, plus minusve linearis, circiter 5 cm. longus et 6 mm. latus, patenter pubescens, nervo marginali et septo membranaceo praeditus. Semina in fructu pleno circiter 12, disciformia, embryone 3 mm. diametro, semina ala hyalina 1.5 mm. lata circumcincta; cotyledones accumbentes 4 mm. longi et 2.5 mm. lati leviter emarginati. ASIA MINOR. Lycia: Vil. Antalya, Distr. Kemer, Takhtali Dağ; 2000-2300 m.; mobile north-facing screes; leaves glaucous fleshy, flowers lilac-pink; 16 Aug. 1947, Davis 14123 (typus, K!); ibid., north scree and north ridge, 10 July 1949, Davis 15096 (K!).

Ricotia varians

Flora of Turkey

R. varians B. L. Burtt in Kew Bull. 1951: 131 (1952). Differs from R. davisiana in being completely glabrous, in its narrowly elliptic to obovate leaf segments, and its 1(-3)-seeded, oval-elliptic fruit. Fl. 8. Screes, 2200-2500 m.

Type: Turkey (C3] vil. Isparta, distr. Sutçuler, Dedegöl Dağ above Obruk Yayla, 2200-2400 m, Davis 15958 (holo. K!).

Endemic, and known only from Dedegöl Da. Ir.-Tur. element. Related to R. davisiana.

1951 Genus Ricotia

GOOGLE TRANSLATE:... from the affinity of R. isatoidei (Barbey) B. L. Burtt from which it is distinguished by larger flowers, fruits sometimes 2-3-sperm (most often monosperm), seeds more broadly winged. R. davisianae B. L. Burtt is also related but easily distinguished by glabrousness and fruits often monosperms never 8-12-sperms. Perennial herb, up to 10 cm. tall, with many erect-flexuous branched stems; entire plant glabrous. Leaves subfleshy, petiolate, trifid or pinnately 4-5-fid lamina attenuated into a cuneate petiole; petiole varying in length, 1-5 cm. long; middle segment of lamina larger than others, narrowly elliptic to obovate, up to 2 cm. long and 1 cm. wide but usually less, apex rounded (sometimes mucronulate) or subacute, usually entire; lateral segments superior but usually smaller and slightly oblique; lower lateral segments (if present) much smaller. Terminal inflorescence ebracteate racemose, on a bare peduncle 6-12 cm. long, 6-15-flowered. Pedicel in anthesis 2.5 mm., under fruit 4 mm. long. Sepals 4, cohering with each other, the exteriors slightly saccate, 3.5 mm. long, 1.5 mm. wide, elliptic, apex slightly hyaline at the margins. Petals pink-lilac, 7 mm. long, claw 3.5 mm. long, lamina obcordate 3.5 mm. long and 4 mm. wide horizontally open. Stamen long filaments 2 mm. long; short filaments 1 mm. long; anthers 1.5 mm. long. Gynoecium 1 mm. barely exceeding, 0.25 mm. wide slightly compressed, stigma sessile. Fruit usually single-spermed, rarely 2-3-spermed, elliptical or obovate-elliptic (immature), about 1.5 mm. [cm presumably?] long and 8 mm. wide. Immature seeds disciform, membranous-winged, 5 mm. in diameter.

ORIG: R. varians B. L. Burtt, sp. nov. ex affinitate R. isatoidei (Barbey) B. L. Burtt a qua floribus majoribus, fructibus interdum 2-3-spermis (saepissime monospermis), semina latius alata distinguitur. R. davisianae B. L. Burtt etiam affinis sed glabritie et fructibus saepissime monospermis nunquam 8-12-spermis facile distinguitur. Herba perennis, 10 cm. usque alta, caulibus pluribus ramosis erecto-flexuosis; tota planta glabra. Folia subcarnosula, petiolata, lamina trifida vel pinnatim 4-5-fida in petiolum cuneatim attenuata; petiolus longitudine varians, 1-5 cm. longus; segmentum laminae medium aliis majus, anguste ellipticum ad obovatum, usque 2 cm. longum et 1 cm. latum sed plerumque minus, apice rotundatum (interdum mucronulatum) vel subacutum, plerumque integrum; segmenta lateralia superiora sed plerumque minora et leviter obliqua; segmenta lateralia inferiora (si adsint) multo minora. Inflorescentia terminalis ebracteata racemosa, pedunculo nudo 6-12 cm. longo 6-15-floro. Pedicellus anthesin 2.5 mm., sub fructu 4 mm. longus. Sepala 4, inter se cohaerentia, exteriores leviter saccata, 3.5 mm. longa, 1.5 mm. lata, elliptica, apice ad margines paullo hyalina. Petala roseo-lilacina, 7 mm. longa, ungue 3.5 mm. longo, lamina obcordata 3.5 mm. longa et 4 mm. lata horizontaliter patente. Stamina longa filamentis 2 mm. longis; brevia filamentis 1 mm. longa; antherae 1.5 mm. longae. Gynoecium 1 mm. vix excedens, 0.25 mm. latum leviter compressum, stigmate sessili. Fructus plerumque mono- spermus, raro 2-3-spermus, elliptica vel obovato-elliptica (immaturus) circiter 1.5 mm [cm presumably?]. longus et 8 mm. latus. Semina immatura disciformia, membranaceo-alata, 5 mm. diametro. ASIA MINOR. Pisidia/Isauria. Prov. Isparta, distr. Sutçuler; Dedegöl Dağ above Oruz Gaz Yayla; mobile south-facing screes; perennial with lilac-pink flowers; 2200-2400 m.; 2 Aug. 1949, Davis 15958 (typus, K!). Ibidem, above the tarn Dedegöl; mobile scree; 2500 m.; 3 Aug. 1949, Davis 16000 (K!).

Ricotia isatoides

2016 Atlas of the Aegean

Glabrous, short-lived perennial, slightly woody at base. Stems slender, suberect, 10-25 cm. Basal and lower cauline leaves broadly oblanceolate, entire to shallowly lobed, glaucous-green or suffused purple. Racemes lax, 10-15-flowered. Petals c. 5 mm, ± violet, truncate. Siliculae pendent, strongly compressed, obovate, 9-12 x 6-8 mm, narrowly winged. Seed usually solitary in each loculus.

1993 Flora Europaea ed 2 vol 1

Fleshy perennial 15-20 cm. Leaves entire, crenate or pinnatifid with 3-5 lobes. Petals c. 5 mm, pale violet or with violet veins. Silicula 10-12 x 6-8 mm, about twice as long as wide, with very narrow marginal wing. Seeds usually solitary. Calsarcous screes. Karpathos, Cr..